FEITOSA DOS SANTOS MACHADO, R. 2011. Revisao taxonomica e analise filogenetica de Heteroponerinae (Hymenoptera, Formicidae). PhD thesis Universidade de Sao Paulo. 311 pages.
HANISCH, P. E.; CALCATERRA, L. A.; LEPONCE, M.; ACHURY, R. ; SUAREZ, V.; SILVA, R. R.; PARIS, C. 2015. Check list of ground-dwelling ant diversity (Hymenoptera: Formicidae) of the Iguazú National Park with a comparison at regional scale. Sociobiology 62(2): 213-227.
HANISCH, P. E.; SUAREZ, V.; TUBARO, P. L.; PARIS, C. I. 2018. Co-occurrence patterns in a subtropical ant community revealed by complementary sampling methodologies. Environmental Entomology 47(6): 1402–1412.